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KYIV, Ukraine (AP) — In a dramatic escalation of East-West tensions over Russia’s invasion of Ukraine, President Vladimir Putin ordered Russian nuclear forces put on high alert Sunday in response to what he called “aggressive statements” by leading NATO powers.

The directive to put Russia’s nuclear weapons in an increased state of readiness for launch raised fears that the crisis could boil over into nuclear warfare, whether by design or mistake.

Putin’s step is “potentially putting in play forces that, if there’s a miscalculation, could make things much, much more dangerous,” said a senior U.S. defense official.

Putin, in giving the nuclear alert directive, cited not only statements by NATO members — who have rushed to reinforce the military alliance’s members in Eastern Europe — but the hard-hitting financial sanctions imposed by the West against Russia, including Putin himself.

Speaking at a meeting with his top officials, Putin told his defense minister and the chief of the military’s General Staff to put nuclear forces in a “special regime of combat duty.”

“Western countries aren’t only taking unfriendly actions against our country in the economic sphere, but top officials from leading NATO members made aggressive statements regarding our country,” Putin said in televised comments.

U.S. defense officials would not disclose their current nuclear alert level except to say that the military is prepared all times to defend its homeland and allies.

White House press secretary Jen Psaki told ABC that Putin is resorting to a pattern he used in the weeks before the invasion, “which is to manufacture threats that don’t exist in order to justify further aggression.”

The practical meaning of Putin’s order was not immediately clear. Russia & the U.S. typically have land & submarine-based nuclear forces on alert and prepared for combat at all times, but nuclear-capable bombers & other aircraft are not.

If Putin is arming or otherwise raising the nuclear combat readiness of his bombers, or if he is ordering more ballistic missile submarines to sea, then the United States might feel compelled to respond in kind, said Hans Kristensen, a nuclear analyst at the Federation of American Scientists. That would mark a worrisome escalation, he said.

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